FANS were horrified to discover Dominic West and Lily James were playing father and daughter in the TV show before being caught kissing in Rome.
The two actors are starring in the Amazon and BBC adaptation of The Pursuit of Love by Nancy Mitford.
Lily, 31, will be Linda Radlett – a “wild and love-addicted” woman hunting down the ideal husband across Europe.
But rather than play her lover, Dominic will be playing her father, called Uncle Matthew, in the TV show.
Fans were horrified by the discovery, especially after pictures of West and James kissing in Rome emerged.
One person said: “Several days behind everyone else on this BUT! The fact that Dominic West and Lily James probably met while filming The Pursuit of Love, where he is playing HER FATHER.”
A second added: “Lily James and Dominic west are working on a show together where they play father and daughter”
A third remarked: “not dominic west playing lily james’ father in their new miniseries. i hate it here. i really do”
The Affair star Dominic was seen getting cosy with his co-star Lily on a break in Rome.
The pair were spotted as they spent two nights in a suite at the top of swanky Hotel De La Ville.
In the pictures Dominic, who married wife Catherine FitzGerald in 2010, is seen stroking Lily’s hair as they get lunch together and smiling at the brunette beauty.
According to insiders, Lily and Dominic grew close after they were “forced into a bubbly” while on set.
A source told The Mail Online: “Due to the pandemic there were strict rules about off-set mingling.
“When you are making films these days, you end up locked down in a bubble for 12 weeks. People go crazy.
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“You aren’t allowed to hang out with people from outside the bubble, so you end up spending time with each other.”
Another source added: “Actors have to sign Covid contracts these days, which mean they promise they will stay in that bubble throughout filming.
“They can’t see their families even at weekends. It means that people are thrown together in a very intense way.”